Download Photosynthesis: The Process of Converting Light Energy into Chemical Energy and more Slides Biology in PDF only on Docsity! Photosynthesis: Capturing Energy Chapter 9 docsity.com Light Consists of particles (photons) that move as waves Photons with shorter wavelengths have more energy than those with longer wavelengths docsity.com Photosynthetic Pigments • In thylakoid membranes – chlorophyll a – chlorophyll b – Carotenoids docsity.com Energizing Electrons • chlorophyll Photons excite photosynthetic pigments • move to electron acceptor compounds Energized electrons docsity.com Active Wavelengths • Combined absorption spectra of chlorophylls a and b – action spectrum for photosynthesis docsity.com The Photosystems Photosystems I and II photosynthetic units include chlorophyll, accessory pigments organized with pigment-binding proteins into antenna complexe docsity.com A Photosystem
Primary
electron
acceptor
Thylakoid
membrane
Photosystem
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LIsNt-aqepenaent
reactions
Primary
electron
SeeePn Electron
transport
chain
Electron
transport
Production
of ATP by
chemiosmosis
0.5
Photosystem | (P70)
Oxidation-reduction potential (volts) (relative energy level)
15
@ Electrons are supplied to system from the splitting of HO by photo- — @ Electrons in photosystem | are “re-energized”
system Il, with release of O as by-product. When photosystem Il is by absorption of additional light energy and
activated by absorbing photons, electrons are passed along electron are passed to NADP", forming NADPH.
transport chain and are eventually donated to photosystem I.
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Cyclic Electron Transport • Electrons from photosystem I – return to photosystem I • ATP produced by chemiosmosis • No NADPH or oxygen generated docsity.com TABLE 9-1
A Comparison of Noncyclic
and Cyclic Electron Transport
Electron source
Oxygen released?
Terminal electron
acceptor
Form in which
energy is temporarily
captured
Photosystem(s)
Cy : : . "
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Noncyclic
Electron
Transport
H,0
Yes (from HO)
NADP*
ATP (by
chemiosmoasis);
NADPH
PS | (P700) and
PS II (P680)
Cyclic
Electron
Transport
None—electrons cycle
through the system
No
None—electrons cycle
through the system
ATP (by chemiosmosis)
PS | (P700) only
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Proton Gradient
* Stroma °°
- . Thylakoid lumen
Thylakoid membrane/ , * Protons (H*)
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Calvin Cycle (C3 pathway) Carbon fixation reactions 3 phases • CO2 uptake (fixation) phase • Carbon reduction phase • RuBP regeneration phase docsity.com CO2 Fixation Phase Enzyme rubisco (ribulose bisphosphate carboxylase/ oxygenase) combines CO2 with ribulose bisphosphate (RuBP), a five-carbon sugar forms 3-carbon phosphoglycerate (PGA) docsity.com Carbon Reduction Phase
Energy of ATP and NADPH
convert PGA molecules to glyceraldehyde-3-
phosphate (G3P)
J
7
For each 6 CO, fixed
2 G3P leave cycle to
produce 1 glucose
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12 G3P are produced
Photorespiration C3 plants use O2 and generate CO2 • by degrading Calvin cycle intermediates • but do not produce ATP On bright, hot, dry days • plants close stomata, conserving water • prevents passage of CO2 into leaf docsity.com C4 Pathway • Takes place in mesophyll cells • Enzyme PEP carboxylase binds CO2 • CO2 fixed in oxaloacetate – converted to malate • Malate moves into bundle sheath cell – CO2 is removed • Released CO2 enters Calvin cycle docsity.com C, and C, Plants
—— . Upper =— ’ i ~
iis ; epidermis : »
Palisade mesophyll
Bundle sheath cells
of veins
(a) In C3 plants, the Calvin cycle takes place in the mesophyll (b) In Cy plants, reactions that fix CO into four-carbon compounds
cells and the bundle sheath cells are nonphotosynthetic. take place in the mesophyll cells. The four-carbon compounds are
transferred from the mesophyll cells to the photosynthetic bundle
sheath cells, where the Calvin cycle takes place.
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